University of Idaho University of Idaho
Cells & Evolution of Life


 

 

  © 2008 All rights reserved.
 
University of Idaho
 
Dept. Biological Sciences
 

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Glossary

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C

C4
Plants that use PEP carboxylase to fix CO2 into a four carbon compound, which is then transferred to special cells, called bundle sheath cells, where the CO2 is fixed by RuBisCo. C4 plants avoid photorespiration by keeping CO2 concentrations high in the vicinity of RuBisCo.

Calvin-Benson cycle
(dark reactions, carbon-fixation reactions)
The reactions by which RuBisCo fixes carbon dioxide to a five carbon molecule (RUBP); the resulting six carbon molecule is quickly split and reduced, ultimately producing carbohydrate.

CAM Plants
Plants that use PEP carboxylase to fix CO2 into a four carbon acid during the night. This acid is stored in the vacuole until daytime, when it is moved out of the vacuole and decarboxylated. The carbon dioxide is then fixed by RuBisCo in the Calvin-Benson cycle. CAM plants avoid photorespiration by building up a store of CO2 during the night, and then using this store during the day to produce carbohydrates and maintain a relatively high CO2 concentration in the vicinity of RuBisCo.

carbohydrate
Organic compounds with the general formula CnH2mOm . Common examples are sugars, starch, and cellulose.

carbon fixation reactions
See Calvin-Benson cycle

carrier proteins
Integral membrane proteins that bind substances in order to move them across a membrane

catabolism
Catabolism encompasses the processes of cell metabolism in which the cell breaks down complex molecules to produce energy and reducing power

catalysis
See catalyst
catalyst
(cat´ a list) [Gr. kata-, implying the breaking down of a compound]
A chemical substance that accelerates a reaction without itself being consumed in the overall course of the reaction. Catalysts lower the activation energy of a reaction. Enzymes are biological catalysts.

cell
The cell is the basic unit of life. All known cells (except a few specialized cell types) have certain basic components in common:

  • cytoplasm (the living substance which makes up most of the cell volume)

  • DNA, the genetic information that acts as a blueprint for the other components.

  • Proteins, the machinery of the cell.

  • Membranes, which separate the cell form its environment, work as a filter, and act as a communication relay with the outside

character
A particular phenotypic feature, such as hair, or skin, or leaf.

chemoautotrophs
See autotroph
 

chloroplast
Chloroplasts are organelles found in plants and eukaryotic algae which conduct photosynthesis. These are one of the forms a plastid may take, and are generally considered to have originated as endosymbiotic cyanobacteria. In green plants they are surrounded by two lipid bilayer membranes, which correspond to the host cell and ancestral bacterium, though algal chloroplasts show notable variations from this pattern. The fluid within the chloroplast is called the stroma, corresponding to the cytoplasm of the bacterium, and contains tiny circular DNA and ribosomes. The genome is considerably reduced compared to that of free-living cyanobacteria, but the parts that are still present show clear similarities.

The structure and biochemistry of chloroplasts are also similar to those of cyanobacteria. Carbon dioxide is converted into carbohydrates via the Calvin cycle, which takes place in the stroma. Thylakoids, tiny membrane-enclose sacs, are present, and in most forms are arranged into stacks called grana. The thylakoid membranes are the site of photosynthesis, converting the energy of light into ATP and oxidizing water to oxygen. The photosynthetic proteins in the membrane bind chlorophyll, which is present with various accessory pigments. These give chloroplasts their colour, which may be green, golden, brown, or red.

chromosome
A chromosome is, minimally, a very long, continuous piece of DNA, which contains many genes, regulatory elements and other intervening nucleotide sequences. In the chromosomes of eukaryotes, the uncondensed DNA exists in a quasi-ordered structure inside the nucleus, where it wraps around histones (structural proteins, Fig. 1), and where this composite material is called chromatin. During mitosis (nuclear division), the chromosomes are condensed and called metaphasic chromosomes. This is the only natural context in which DNA is visible with an optical microscope. Prokaryotes do not possess histones or nuclei. In its relaxed state, the DNA can be accessed for transcription, regulation, and replication.

Autosomes are chromosomes that carry genes not related to sex determination; sex chromosomes carry genes related to the determinance of sexual characteristics.

Citric acid cycle
In biochemistry, the Citric Acid Cycle, also called Krebs cycle, Citrate Cycle, Tricarboxylic Acid Cycle or TCA, is a series of chemical reactions of central importance in all living cells that utilize oxygen as part of cellular respiration. In these aerobic organisms, the citric acid cycle is a metabolic pathway that forms part of the break down of carbohydrates, fats and proteins into carbon dioxide and water in order to generate energy.

The Krebs cycle is named after Sir Hans Adolf Krebs (1900-1981), who was awarded the 1953 Nobel Prize in Medicine for its discovery.

The citric acid cycle takes place within the mitochondria in eukaryotes, and within the cytoplasm in prokaryotes.

The citric acid cycle forms part of carbohydrate catabolism, protein catabolism and fat catabolism. All these three processes produce acetyl-CoA, a two-carbon acetyl group bound to coenzyme A. Acetyl-CoA is the main input to the citric acid cycle.

cleavage
An early embryonic stage of development during which the zygote divides to produce two, four, and then eight cells.

clone
Genetically identical cells or organisms. Cells may produce clones by fission (prokaryotes) or mitosis (eukaryotes. Some multicellular organisms produce clones by asexual reproduction or by vegetative propagation.

codominance
A phenomenon whereby both alleles of a gene in a diploid organism are expressed and active in the phenotype (ex. blood types of humans).

codon
A codon is a sequence of three bases in RNA.  Each codon specifies one amino acid. For example, the RNA sequence UUUAAACCC specifies three codons (UUU-AAA-CCC), which each specify one amino acid. This RNA sequence, then, encodes a protein sequence three amino acids in length (as we will see, it encodes Phenylalanine-Lysine-Proline). There are sixty-four possible codons.

Nearly all living things use the same genetic code. The standard version is given in the following tables, which show what amino acid each of the 43 = 64 possible codons specify (Table 1), and what codons specify each of the 20 amino acids involved in translation.  For instance, GAU codes for the amino acid Asp (asparagine), and Cys (cysteine) is coded for by the codons UGU and UGC.  The bases in the table below are adenine, cytosine, guanine and uracil, which are used in the mRNA; in the DNA, thymine takes the place of uracil.

Table 1 : Codon table. This table illustrates the 64 possible codon triplets.

  2nd base
U C A G
1st base U UUU Phenylalanine
UUC Phenylalanine
UUA Leucine
UUG Leucine
 
UCU Serine
UCC Serine
UCA Serine
UCG Serine
 
UAU Tyrosine
UAC Tyrosine
UAA Ochre Stop
UAG Amber Stop
 
UGU Cysteine
UGC Cysteine
UGA Opal Stop
UGG Tryptophan
 
C CUU Leucine
CUC Leucine
CUA Leucine
CUG Leucine
 
CCU Proline
CCC Proline
CCA Proline
CCG Proline
 
CAU Histidine
CAC Histidine
CAA Glutamine
CAG Glutamine
 
CGU Arginine
CGC Arginine
CGA Arginine
CGG Arginine
 
A AUU Isoleucine
AUC Isoleucine
AUA Isoleucine
1AUG Methionine
 
ACU Threonine
ACC Threonine
ACA Threonine
ACG Threonine
 
AAU Asparagine
AAC Asparagine
AAA Lysine
AAG Lysine
 
AGU Serine
AGC Serine
AGA Arginine
AGG Arginine
 
G GUU Valine
GUC Valine
GUA Valine
GUG Valine
 
GCU Alanine
GCC Alanine
GCA Alanine
GCG Alanine
 
GAU Aspartic acid
GAC Aspartic acid
GAA Glutamic acid
GAG Glutamic acid
 
GGU Glycine
GGC Glycine
GGA Glycine
GGG Glycine
 

1The AUG codon both codes for methionine and serves as an initiation site; the first AUG in an mRNA's coding region will be the site where translation into protein begins.

 

coenzymes
Small carbon molecules that bind to and are required for the activity of certain enzymes.

cofactors
Molecules essential for the activity of some enzymes; these can be inorganic ions or organic molecules.

competitive inhibitors
Inhibition of an enzyme where the substrate and inhibitor compete for the active site(s).

complementary base-pairing
The pairing of complementary nitrogenous bases between the two strands of the DNA molecule, between DNA and RNA, or between different segments of an RNA molecule. The pairings are as follows: adenine (A) pairs with either thymine (T) or uracil (U), while guanine pairs with cytosine.

condensation reaction
Also see dehydration synthesis
Common reaction of biosynthetic pathways, such as those that build macromolecules. During this reaction, two monomers are covalently linked, with the concurrent release of a molecule of water. The water forms from the combination of the hydroxyl (OH) group of one monomer and a hydrogen from the other monomer.

conjugation
Need Definition...

covalent bond
Covalent bonding is a form of chemical bonding characterized by the sharing of one or more pairs of electrons by two atoms to produce a mutual attraction. The atoms tend to share electrons, so as to fill their outer electron shells. Commonly covalent bond implies the sharing of just a single pair of electrons. The sharing of two pairs is called a double bond and three pairs is called a triple bond.  Covalent bonding tends to be stronger than other types of bonding.

crossover
Crossover is the process by which two chromosomes paired up during prophase I of meiosis exchange a distal portion of their DNA. Crossover occurs when two chromosomes, normally two homologous instances of the same chromosome, break and connect to each other's ends. If they break at the same locus, this merely results in an exchange of genes. This is the normal way in which crossover occurs. If they break at different loci, the result is a duplication of genes on one chromosome and a deletion on the other. If they break on opposite sides of the centromere, this results in one chromosome being lost during cell division.

Any pair of homologous chromosomes may be expected to cross over three or four times during meiosis. This aids evolution by increasing independent assortment, and reducing the genetic linkage between genes on the same chromosome.

cytokinesis
Cytokinesis refers to the division of a eukaryotic cell. Cytokinesis generally follows the replication of the cell's chromosomes, usually mitotically, but sometimes meiotically. Except for some special cases, the amount of cytoplasm in each daughter cell is the same. In animal cells, the cell membrane forms a cleavage furrow and pinches apart like a balloon. In plant cells, a cell plate forms, which becomes the new cell wall separating the daughters. Various patterns occur in other groups.

cytoplasm
The cytoplasm refers to everything bound by the plasma membrane except the nucleus. Perhaps a more useful distinction would be to talk about the cytosol. The cytosol is a watery matrix, bound by the plasma membrane, which contains the cell's organelles and other components. Small molecules, dissolved ions, and soluble macromolecules can also be found in the cytoplasm.